Ery obscurely tuberculate. Flowers about 84 per capitulum, the pistillate flowers peripheral, the staminate central, equalling to subequalling the pistillate in number; the receptacle sparingly pilose or pilose only toward center. Receptacular bracts equaling to subequaling flowers, broadly linear-subspatulate, ca. 6 occasions longer than wide, the apex often MedChemExpress NAMI-A slightly cucullate and pubescent as the sepals, the base distinctly carinate, clasping. Pistillate flowers: Pedicels ca. 0.1.35 mm, thick, glabrous, often becoming callose-thickened in fruit (Peru), leaving characteristic `stumps’ around the empty receptacle following abscission of flowers. Sepals broadly obovate, strongly cymbiform, with apex convex-acute to obtuse (1.35 1.five.0 mm long 0.6.0 mm wide (width variable within a flower), (0.25 0.35.4 (.five) mm wide at the base; black-mottled on shoulders, the midvein location paler brown; shortciliate along upper margin and bearded with longer appressed hairs on upper dorsum; membranaceous to chartaceous at anthesis, enlarging slightly and becoming uniformly thickened, and typically rigid in fruit, husk-like and non-hygroscopic, enclosing the corolla and fruit and dispersed with it. Petals oblanceolate-spatulate, obtuse to usually emarginate or truncate-emarginate, (1.2 1.35.7 (.85) mm lengthy (.35 0.40.7 (.8) mm wide, ca. 2.two.two (.9) occasions longer than wide, cream to brownishtinged, with scale-like staminode at base, pilose abaxially near margins of distal half with much more or significantly less tuberculate trichomes, also densely tufted subapically within in two patches either side of your apex, which enfold the style branch; like the sepals becoming a lot more or less rigid-thickened in fruit, the staminode also thickening and tightly adherent to both petal and ovary base. Gynoecium with ovary 0.5 mm at anthesis, ca. 0.85 mm in fruit; the style base 0.3.five mm lengthy; the nectaries with stalks 0.25.65 mm PubMed ID:http://www.ncbi.nlm.nih.gov/pubmed/20107779 lengthy, glandular portion 0.2.35 mm lengthy, quite dark brown or dark reddish, subclavate, with a ring of brown membranous papillae ca. 2 rows thick in the apex, the whole structure keeping its shape after anthesis; style branches 0.6.0 (.two) mm extended, ordinarily ca. 0.2 mm longer than nectaries though frequently establishing unequally, the stigmas simple, dark red-brown, non-involute right after anthesis. Seeds subglobose to ellipsoid, largely 0.665 mm long, 0.4.55 mm wide, red-brown, reticulate with quick weak pseudotrichomes, at times glabrate. Staminate flowers: Pedicels 0.15 mm to obsolete. Sepals (1.two 1.35.75 mm extended 0.five.85 mm, commonly strongly and unequally fused at base for 3/4 their length, obovate-navicular above, the apex obtuse-angled to broadly rounded, the tubular base usually rigid and obconic at maturity; color and pubescence as within the pistillate flowers. Corolla including anthophore 1.35.9 mm; the anthophore 0.65.35 mm long, generally ca. 600 the corollaNancy Hensold / PhytoKeys 64: 17 (2016)Figure four. Habit of Paepalanthus caryonauta A Boyle 4219 B Fuentes 15374 C-D Valenzuela 8117.length, ca. 0.two.35 mm diameter at base, fleshy and columnar at maturity; the corolla tube 0.35.75 mm deep, fleshy towards base specially opposite the filaments, the corolla lobes hyaline to brownish, obtuse, 0.15.35 mm, not involute soon after anthesis; intermediate lobes lacking. Stamen filaments with the basal half fleshy, terete and adnate to the corolla, abruptly narrowed and loosely adhering to the lobes above, exsert 0.two.five mm beyond the lobes, the exsert portion dark reddish-brown specifically at tip; anthers cream-c.